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Kukushkin O. V., Doronin I. V., Tuniyev B. S., Ananjeva N. B., Doronina M. A. INTRODUCTION OF AMPHIBIANS AND REPTILES IN THE CAUCASUS AND CRIMEA: AN OVERVIEW AND SOME ACTUAL DATA. Current Studies in Herpetology, 2017, vol. 17, iss. , pp. 157-?. DOI:

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An overview of the cases of introduction (both accidental and intentional) of amphibians and reptiles in the Caucasus and Crimea is provided. The introduction cases are systemized in accordance with the goals and ways of introduction and with regards to the expected scale of the impact and its nature. No negative consequences of the introduction of amphibian and reptile species in the Crimea and Caucasus are currently observed on any of the known examples. No cases of species naturalization, which the “invasion” term can be applied to, have been revealed. The importance of allochthonous populations as experimental sites to study microevolutionary processes and ecological adaptations of the species in new environmental conditions is discussed.


Alekperov A. M. Changes in the Herpetofauna of the Absheron Peninsula and Theirs Main Causes. Scientific Notes of Azerbaijan State University, Series Biological Sciences, 1973, no. 4, pp. 46–49 (in Russian).
Alekperov A. M., Bogdanov O. P. Reptiles as Inhabitants of the Sheep-farms. Scientific Notes of the Azerbaijan State University, Series Biological Sciences, 1979, no. 2, pp. 44–45 (in Russian).
Alekperov A. M., Galaeva N. M. Morphology of a Secret Toadhead Agama, Acclimatized on the Absheron. Scientific Notes of Azerbaijan State University, Series of Biological Sciences, 1974, no. 2, pp. 63–66 (in Russian).
Alekperov A. M., Loginov А. А. Essays on the biology of reptiles of the Absheron. Proc. of Azerbaijan State University, Series Biological, 1953, vol. 5, pp. 106–127 (in Russian).
Alekperov A. M., Gazanchan M. K., Mamedova S. A. About a finding of the Secret Toadhead Agama in Azerbaijan. In: Herpetology. Tashkent, Nauka, 1965, p. 102. (in Russian).
Ananjeva N. B., Orlova V. F. Distribution and Geographic Variability of Caucasian Rock Agama, Agama caucasia (Eichwald, 1831). Proc. of the Zoological Institute of Academy of Sciences of USSR, vol. 89: Ecology and Systematics of Amphibians and Reptiles. Leningrad, 1979, pp. 4–17 (in Russian).
Ananjeva N. B., Orlov N. L., Khalikov R. G., Darevsky I. S., Ryabov S. A., Barabanov A. V. The Reptiles of Northern Eurasia (Taxonomic Diversity, Distribution, Conservation Status). Saint Petersburg, Zoological Institute of the Russian Academy of Sciences, 2006. 232 p. (in Russian).
Andreev V. Yu. Caspian Bent-Toed Gecko – Tenuidactylus (Cyrtopodion) caspius (Eichwald, 1831). Red Data Book of the Astrakhan Region. Rare and endan-gered objects of flora and fauna. 2nd edition. Astrakhan, “Astrakhan University”, 2014, pp. 266–267 (in Russian).
Antipina E. V., Maslov S. P. History of the formation of ecosystems of the Crimea in the Late Holocene. In: Abstracts of All-Union Zoogeographical Conference. Moscow, 1984, pp. 164–165 (in Russian).
Baskakova M. V. Phylogenetic position of the Green lizard from the Crimea. Biology: from the Molecule to the Biosphere: Abstracts of 10th International Conference of Young Scientists. Kharkiv, T. N. Shapovalov Publ. House, 2015, pp. 169 (in Russian).
Belik V. P. On Amphibian Fauna and Ecology in the Steppe part of the Don Basin. Current Studies in Herpetology, 2010, vol. 10, iss. 3–4, pp. 89 – 100 (in Russian).
Beskaravaynyi M. M., Kotelnikov S. N. The Results of the Gecko Cyrtopodion kotschyi Introduction in the Karadag Nature Reserve. Vestnik Zoologii, 2001, vol. 35, no. 1, pp. 53 (in Russian).
Biological invasions in aquatic and terrestrial ecosystems. Moscow, KMK Scientific Press Ltd., 2004. 436 p. (in Russian).
Bobrov V. V., Warshavsky A. A., Khlyap L. A. Alien Species of Mammals in the Ecosystems of Russia. Moscow, KMK Scientific Press Ltd., 2008. 232 p. (in Russian).
Bunyatova S. N., Akhmedov S. B., Jafarov A. R. Ecological analysis of reptiles (Testudinidae, Sauria, Serpentes) of Talysh. Bull. of the Samara Scientific Center of the Russian Academy of Sciences, 2012, vol. 14, no. 1, pp. 144 – 149 (in Russian).
Vereshchagin N. K. Mammals of the Caucasus. Moscow ; Leningrad, Publ. House of Academy of Sciences of USSR, 1959. 704 p. (in Russian).
Vereshchagin N. K. Experience of the Lizards Ressettlement. Priroda, 1966, no. 11, pp. 75–77 (in Russian).
Vereshchagin N. K. From the Muskrat to the Mammoth. The way of the Zoologist. Saint Petersburg, Asterion, 2002. 336 p. (in Russian).
Vereshchagin N. K. My Century. Memories and Scientific Works. Tver’, A Seventh Letter, 2008. 244 p. (in Russian).
Gorbunov R. V, Gorbunova T. Y., Kononova N. K. Climatic norms of an air temperature on the territory of the Crimea peninsula. Culture of the Black Sea Region Peoples, 2014, vol. 2, no. 278, pp. 89–94 (in Russian).
Dal’ S. K. Landscape-ecological essay of vertebrate animals from places of possible natural foci of the brucellosis in the Stavropol region. Proc. of the Caucasus and Transcaucasia Anti-Plague Research Institute. Stavropol’, 1959, iss. 2, pp. 93–114 (in Russian).
Darevsky I. S. Study and protection of the rare and endangered amphibians and reptiles species of the USSR’ fauna. Study and Protection of the Rare and Endangered Animals of the Fauna of USSR. Moscow, Nauka, 1985, pp. 37–43 (in Russian). 
Darevsky I. S. Consequences of the Failed Attempt to Introduce of the Bisexual Species of the Rock Lizard Darevskia mixta (Mehely, 1909) (Sauria, Lacertidae) from Georgia to the Zhytomyr Region of Ukraine. Vestnik Zoologii, 2006, vol. 40, no. 4, pp. 370 (in Russian).
Darevsky I. S., Orlov N. L. Rare and Endangered Animals. Amphibians and Reptiles. Moscow, Vyshaya shkola, 1988. 463 p. (in Russian).
Darevsky I. S., Szczerbak N. N. Acclimatization of Parthenogenetic Lizards in the Ukraine. Priroda, 1968, no. 5, p. 93 (in Russian).
Dbar R. S., Malandzia V. I. Gymnodactylus caspius Eichw. (Reptilia. Gekkonidae) in Abkhazia. Proc. of the 2nd Regional Conference “Biological Diversity of the Caucasus”. Sukhum, Abkhazian State University Press, 2002, pp. 95–97 (in Russian).
Deryugin K. M. A Journey and Zoological Studies in Coruh River Area (South-Western Transcaucasia) and suburbs of Trapezund. Proc. of Imperial Saint Petersburg Society of Naturalists. Department of Zoology and Physiology, 1899, vol. 30, iss. 2, pp. 49–115 (in Russian).
Doronin I. V. The use of GIS for the Analysis of the Distribution of Rock lizards Darevskia (saxicola) complex (Sauria: Lacertidae). Current Studies in Herpetology, 2012, vol. 12, iss. 3–4, pp. 91–122 (in Russian). 
Doronin I. V. New Records of Rock lizards of the Genus Darevskia Arribas, 1997 (Sauria: Lacertidae) in the Caucasus. Proc. of the Zoological Institute of the Russian Academy of Sciences, 2013, vol. 317. no. 3, pp. 282– 291 (in Russian).
Dotsenko I. B., Darevsky I. S. A Record of the Darevskia dahli in Darevskia armeniaca Population, In-troduced into the Territory of Ukraine. Proc. of the First Conference of the Ukrainian Herpetological Society. Kyiv, Zoomuseum NMNH NAS of Ukraine, 2005, pp. 47–50 (in Russian).
Dotsenko I. B., Peskov V. N., Miropolskaya M. V. Comparative analysis of external morphology of parthenogenetic and bisexual Rock lizards of a genus Darevskia from the territory of Ukraine, and the question of belonging to the species of doubtful introduced lizards specimens. Proc. of Zoological Museum NMNH NAS of Ukraine. Kyiv, 2009, no. 40, pp. 130–142 (in Russian).
Duz’ S. L., Kukushkin O. V., Nazarov R. A. A Record of the Turkestan Nakedtoed Gecko, Tenuidactylus fedtschenkoi (Sauria: Gekkonidae) in the South-Western Ukraine. Current Studies in Herpetology, 2012, vol. 12, iss. 3–4, pp. 123–133 (in Russian).
Dunayev Е. А., Orlova V. F. Class Reptilia. Types of vertebrates in the Zoological Museum of Moscow University. Archives of the Zoological Museum of Moscow State University, 2001, vol. 41, pp. 92–98 (in Russian).
Zhdokova M. K., Shlyakhtin G. V., Zavialov Y. V. Herpetofauna of Kalmykia: species composition, relative abundance, inter-centenery dynamics of distribution. Povolzhskiy J. of Ecology, 2002, iss. 2, pp. 158–162 (in Russian).
Zinenko O. I., Goncharenko L. A. Catalogue of Collections of the Museum of Nature at V. N. Karazin’s Kharkiv National University. Reptiles (Reptilia): Rhynchocephalia; Squamata: Lizards (Sauria), Amphisbaenians (Amphisbaenia). Kharkiv, V. N. Karazin Kharkiv National University Press, 2011. 100 p. (in Russian).
Iljukh M. P. Redeared Slider Trachemys scripta as a New Species of Herpetofauna of Stavropol Region. Science. Innovation. Technologies, 2015, no. 1, pp. 122–126 (in Russian).
Inozemtsev A A., Pereshkolnik S. L. Save a Vestige of the Ancient Pontida. Priroda, 1987, no. 8, pp. 38–49 (in Russian).
Kessler К. F. Journey with the Zoological Purpose to the Northern Shore of the Black Sea and the Crimea in 1858. Kiev, University Press, 1860. 248 p. (in Russian).
Kidov A. A., Matushkina K. A., Timoshina A. L. New Distribution's Materials for Amphibians and Reptiles in Talysh Mountains and Lenkoran Lowland: Some results of Herpetological Expeditions, 2009 – 2011. Proc. of the Ukrainian Herpetological Society, 2011, no. 3, pp. 56–63 (in Russian).
Kidov A. A., Matushkina K. A., Timoshina A. L. On the Habitat Expansion of Caspian bent-toed gecko (Cyrtopodion caspius (Eichwald, 1831)) and Caucasian cat snake (Telescopus fallax (Fleischmann, 1831)) in the Southeastern Azerbaijan. Current Studies in Herpetology, 2012, vol. 12, iss. 1–2, pp. 56–60 (in Russian).
Kidov A. A., Matushkina K. A., Afrin К. А. The first results of captive breeding and reintroduction of the Karelin’s newt, Triturus karelinii Strauch, 1870, from Talysh population. Bull. of the Buryat State University, 2015, iss. 4–1, pp. 81–89 (in Russian).
Kotenko T. I. Protection of Amphibians and Rep-tiles in Nature Reserves of Ukraine. In: Amphibians and Reptiles of Reserved Areas: Collection of Scientific Pa-pers. Moscow, Regional Printing-House of Kalinin city, 1987, pp. 60–80 (in Russian). 
Kotenko T. I. Examples of fluctuations of the spatial allocation in amphibians and reptiles at the south of the Ukraine. Proc. of the First Conference of the Ukrainian Herpetological Society. Kyiv, Zoomuseum NMNH NAS of Ukraine, 2005, pp. 71–75 (in Russian).
Kotenko T. I. On the Distribution of Steppe Viper, Vipera renardi (Reptilia, Viperidae), in the Western part of the Crimean Plain. Vestnik Zoologii, 2007, vol. 41, no. 5, pp. 422 (in Russian).
Kotenko T. I. Amphibians and Reptiles of the Crimea. Scientific Notes of the “Cape Martyan” Nature Reserve. Yalta, 2010, no. 1, pp. 171–224 (in Russian).
Kotenko T. I., Kukushkin O. V. Annotated lists of amphibians and reptiles of the Crimean nature reserves. Scientific Notes of the “Cape Martyan” Nature Reserve. Yalta, 2010, no. 1, pp. 225–261 (in Russian).
Kuzmin S. L. The Amphibians of the Former USSR. Moscow, KMK Scientific Press Ltd., 1999. 298 p. (in Russian).
Kukushkin O. V. Distribution, habitat allocation and abundance of the Kotschy’s (Crimean) gecko, Cyrtopodion kotschyi danilewskii (Strauch, 1887) (Reptilia: Squamata: Gekkonidae) in the Southern Crimea. Karadag. History, Geology, Botany, Zoology: Collection of Scientific Papers. Simferopol’, SONAT, 2004, book 1, pp. 367–396 (in Russian).
Kukushkin O. V. A found of the large exoantropic population of Kotschy’s gecko, Mediodactylus kotschyi danilewskii (Strauch, 1887) (Reptila, Sauria, Gekkoni-dae), on the Crimean South-Eastern Coast. Proc. of the First Conference of the Ukrainian Herpetological Society. Kyiv, Zoomuseum NMNH NAS of Ukraine, 2005, pp. 83–86 (in Russian).
Kukushkin O. V. About inhabitation of Kotschy’s naked-toed gecko, Mediodactylus kotschyi danilewskii (Reptila: Sauria: Gekkonidae), in the middle forest belt of southern macroslope of the Crimean Mountains. Proc. of Ukrainian Herpetological Society, 2009 a, no. 2, pp. 27–36 (in Russian).
Kukushkin O. V. About some patterns of spatial Distribution of Lindholm’s Rock Lizard Darevskia lindholmi (Sauria, Lacertidae) in the South-Eastern coast of the Crimea. Samarskaya Luka: Issues of Regional and Global Ecology, 2009 b, vol. 18, no. 1, pp. 68–75 (in Russian).
Kukushkin O. V. Genezis gerpetofauny Kryma: novoe videnie problemy [Genesis of the Crimean Herpetofauna: a New Vision of the Problem]. In: Materialy nauch. konf. “Iubileinye zoologicheskie chteniia” [Proc. of the Intern. Scientific Conference Devoted to 100th anniversary of S. L. Delyamure and 90th anniversary of S. A. Skryabin “Commemorative Zoological Readings”]. Simferopol’, 2013 a, pp. 22–25 (in Russian).
Kukushkin O. V. Adventivnaia gerpetofauna Ukrainy [Adventive Herpetofauna of the Ukraine]. In: Materialy nauch. konf. “Iubileinye zoologicheskie chteniia” [Proc. of the Intern. Scientific Conference Devoted to 100th anniversary of S. L. Delyamure and 90th anniversary of S. A. Skryabin “Commemorative Zoological Readings”]. Simferopol’, 2013 b, pp. 25–27 (in Russian).
Kukushkin O. V. European pond turtle Emys orbicularis (Linnaeus, 1758). In: Red Data Book of Republic of the Crimea. Animals. Simferopol’, Arial, 2015 а, p. 289 (in Russian).
Kukushkin O. V. Kotschy’s gecko Mediodactylus kotschyi (Steindachner, 1870). In: Red Data Book of Republic of the Crimea. Animals. Simferopol’, Arial, 2015 b, p. 290 (in Russian).
Kukushkin O. V. Giant glass lizard Pseudopus apodus (Pallas, 1775). In: Red Data Book of Republic of the Crimea. Animals. Simferopol’, Arial, 2015 c, p. 291 (in Russian).
Kukushkin O. V., Doronin I. V. Experience of the employment of MaxEnt software for analysis of distributional ranges of the Crimean amphibians and reptiles. Communication 2: Complexities of the data interpretation. Programm and Abstracts of Intenational Scientific Conference Dedicated for 50th Anniversary of the Zoological Museum of Taurida Academy of V. I. Vernadsky Crimean Fed-eral University. Simferopol’, 2015, pp. 63–65 (in Russian).
Kukushkin О. V., Kotenko T. I. Giant legless lizard, sheltopusik Pseudopus apodus (Pallas, 1775). Red Data Book of the Ukraine. Animal kingdom. Kyiv, Publ. House “Globalconsulting”, 2009, pp. 388 (in Ukrainian).
Kukushkin O. V., Tsvelykh A. N. Distribution and Ecological-morphological Peculiarities of the Leopard Snake, Elaphe situla (Serpentes, Colubridae), in the Crimea. Zoologicheskii zhurnal, 2004, vol. 83, iss. 4, pp. 439–448 (in Russian).
Kukushkin O. V., Shaganov V. V. Survey of the Herpetofauna of Opuk Nature Reserve: List of the Species, Theirs Spatial Allocation, and Recommendations for the Protection. Nature Reserves of the Crimea – 2007: Proceedings of 4th International Scientific-practical Conference. Simferopol’, 2007, pp. 93–103 (in Russian).
Kukushkin О. V., Petrov B. P., Nazarov R. A., Melnikov D. A. The Problem of a Biogeographical Status of two Reptiles Species with Narrow Distribution in the Crimean Mountains and the Importance of Cape Aiya Karst Caves for its Solution. Biospelaeological Studies in Russia and Adjacent Countries: Proceedings of 2nd All-Russian Conference of Young Scientists. Yaroslavl’, Filigran’, 2017, pp. 56–69 (in Russian).
Кurtyak F. F., Kurtyak M. F. Red-eared slider Trachemys scripta elegans (Wied, 1839) (Reptilia; Testu-dines), as an Invasive threat in Transcarpathian Region. Scientific Bull. of Uzhgorod University, Ser. Biology, 2013, iss. 34, pp. 58–62 (in Ukrainian). 
Leontyeva O. A., Sichevskij Je. A., Kolonin G. V. Dynamic of the Number of Ticks Hyalomma aegyptium (Ixodidae), Parasite of Mediterranean Tortoise Testudo graeca nikolskii in the Peninsula Abrau (Northwest Caucasus). Bull. of Moscow Society of Naturalists, Ser. Biological, 2016, vol. 121, iss. 1, pp. 26–34 (in Russian).
Litvinchuk S. N. Distribution and Conservation Status of the Banded Newt, Ommatotriton ophryticus (Amphibia: Caudata). Nature Conservation Research, 2017, vol. 2, no 1, pp. 33–39 (in Russian).
Lotiev K. Yu. Lizards (Sauria) in the Red Book of Chechen Republic. Samarskaya Luka: Problems of Regional and Global Ecology, 2009, vol. 18, no. 1, pp. 91–95 (in Russian).
Lyapkov S. M., Ermakov O. A., Titov S. V. Distribution and Origin of Two Forms of the Marsh Frog Pelophylax ridibundus complex (Anura, Ranidae) from Kamchatka, Based on Mitochondrial and Nuclear DNA data. Zoologicheskii zhurnal, 2017, vol. 96, no. 11, pp. 1384–1391 (in Russian).  
Mazanaeva L. F. Caspian gecko – Cyrtopodion caspius (Eichwald, 1831). The Red Book of the Republic of Dagestan. Makhachkala, Ministry of Natural Resources and Environmental Protection of the Republic of Dagestan, 2009, pp. 385–386 (in Russian).
Mazanaeva L. F., Orlova V. F. New Records of Lizards (Sauria: Lacertidae, Scincidae) in Dagestan. Bull. of Moscow Society of Naturalists, Ser. Biological, 2009, vol. 114, no. 4, pp. 63–66 (in Russian).
Mazanaeva L. F., Chernaya A. R. New data on the biology and distribution of the Caspian gecko in Dagestan. Materials of the 14th Scientific-practical Conference on the Protection of the Nature of Dagestan. Makhachkala, 1997, pp. 68–69 (in Russian).
Matvyeyev A. S., Kukushkin O. V., Sokolov L. V. Commom Wall Lizard, Podarcis muralis (Sauria, Lacertidae), as a New Species in the Fauna of Ukraine. Proc. of Ukrainian Herpetological Society, 2013, no. 4, pp. 95–108 (in Russian).
Nazarov R. A., Poyarkov N. A. A Taxonomic Re-vision of the Genus Tenuidactylus Szczerbak et Golubev 1984 (Reptilia, Squamata, Gekkonidae) with a Description of a New Species from Central Asia. Zoologicheskii zhurnal, 2013, vol. 92, no. 11, pp. 1312–1332 (in Russian).
Nekrasova O. D., Kolosh T. V., Bolotov M. B. Significance of Protection of Odessa Lagoon Ecosystems of Black Sea Region on the Example of Herpetological Assemblages. Biodiversity and Role of the Animals in Ecosystems: Materials of the 7th Intern. Conference. Dnepropetrovsk, Adverta, 2013, pp. 234–235 (in Russian).
Nesterov P. V. Materials on the herpetology of the south-western Transcaucasia (Coruh area). Annual Report of Zoological Museum of Imperial Academy of Scienses, 1911, vol. 16, no. 1, pp. 1–18 (in Russian).
Nikitenko M. F. Reptiles of the Soviet Bukovina. In: Animal Kingdom of the Soviet Bukovina. Chernovtsy, Chernovtsy University Press, 1959, pp. 134–160 (in Russian).
Nikolsky A. M. Vertebrate Animals of the Crimea. Scientific Notes of Imperial Academy of Sciences. Physical-Mathematical Branch, 1891, vol. 68, suppl. 4. 484 p. (in Russian).
Nikolsky A. M. Reptiles (Reptilia). Vol. 1. Chelonia et Sauria. Fauna of Russia and Adjacent Countries, Chiefly by Collections of Zoological Museum of Imperial Academy of Sciences. Petrograd: Printing-House of Imperial Academy of Sciences, 1915. 534 p. (in Russian).
Nikolsky A. M. Reptiles (Reptilia). Vol. 2. Ophidia. Fauna of Russia and Adjacent Countries, Chiefly by Collections of Zoological Museum of Imperial Academy of Sciences. Petrograd, Printing-House of Imperial Academy of Sciences, 1916. 350 p. + 8 tables with illustrations (in Russian).
Omelchenko A. V., Girnyk А. Е., Osipov F. А., Petrosyan V. G., Vergun А. А., Ryskov А. P. Detection of Genotypic Changes in the Parthenogenetic Lizards of Darevskia armeniaca (Mehely) Introduced from Armenia to Ukraine. Russ. J. of Biological Invasions, 2016, no. 2, pp. 102–115 (in Russian).
Pestov M. V., Kalinina O. N., Grankina L. I. Cas-pian Benttoed Gecko – Cyrtopodion caspius (Sauria, Gekkonidae) – a New Species in Astrakhan Region and the Volga basin. Samarskaya Luka: Problems of Regional and Global Ecology, 2009, vol. 18, no. 1, pp. 108–110 (in Russian).
Pidoplichko I. G. About the glacial period. Iss. 2. Biological and Geographical Peculiarities of European Representatives of Quaternary Fauna. Kiev, Publ. House of Academy of Sciences of Ukrainian SSR, 1951. 265 p. (in Russian).
Pidoplichko I. G. About the Glacial Period. Iss. 3. History of the Quaternary Fauna of European part of the USSR. Kiev, Publ. House of Academy of Sciences of Ukrainian SSR, 1954. 221 p. (in Russian).
Pisanets E. M. Catalogue of Collection of Zoological Museum NMNH, NAS of Ukraine. The Tailed Amphibians (Amphibia: Caudata). Kyiv, Zoomuseum NMNH NAS of Ukraine, 2003. 148 p. (in Russian).
Pysanets E., Kukushkin O. Amphibians of the Crimea. Kyiv, National Museum of Natural History, 2016. 320 p.
Popov V. N. Amphibians and Reptiles. In: 47th Territory of Priority. Sasyk-Sivash Lake. Simferopol’, Association of Support for Biological and Landscape Diversity in the Crimea – “Gurzuf – 97”, 2000, pp. 19–20 (in Russian).
A Sand Lizard. Monographic description of the species. Ed. A. V. Yablokov. Moscow, Nauka, 1976. 376 p. (in Russian).
Puzanov I. I. Zoogeography. Moscow, Educational-Pedagogical Edition of People Comissariat of Education of Russian SFSR, 1938. 359 p. (in Russian).
Puzanov I. I. Originality of the Crimean Fauna and its Origin. Scientific Notes of Gor’kovskyi State University, 1949, iss. 14, pp. 5–32 (in Russian).
Puzanov I. I. Animal kingdom of Karadag. In: The Karadag. Popular-science essays. Kiev, Publ. House of Academy of Sciences of Ukrainian SSR, 1959, pp. 25–39 (in Russian).
Puzanov I. I. Around the Untrodden Unknown Crimea. Moscow, State Edition of Geogrephical Literature, 1960. 284 p. (in Russian).
A Steppe Rucerrunner. Ed. N. N. Szczerbak. Kiev, Naukova dumka, 1993. 238 p. (in Russian).
Rashkevich N. A. A Snake-eyed Lizard in Checheno-Ingushetia. Priroda, 1975, no. 6, p. 94 (in Russian).
Rashkevich N. A., Semenikhina T. A. Data on Ecology of the Lizards of Checheno-Ingushetia. Bull. of the North Caucuasus Scientific Center of Higher Educational School, Ser. Natural-Historical – Geographical, 1974, no. 3, pp. 98–101 (in Russian).
Sarayev F. A., Pestov M. V. To the Cadastre of Reptiles of Northern and North-Eastern Caspian Sea Region. In: Herpetological Studies in Kazakhstan and Adjacent Countries: Collection of Scientific Papers. Almaty, 2010, pp. 172–191 (in Russian).
Semyonov [Tyan-Shanskyi] A. Several considerations about the past of the Crimean fauna and flora in relation with a record here of a Mountain partridge (Caccalus chukar G. R. Gray). Scientific Notes of Imperial Academy of Sciences, Physical-Mathematical Branch, 1899, vol. 8, no. 6, pp. 3 –19 (in Russian).
Semyonov-Tyan-Shanskyi A. Limits and Zoogeographical Subdivisions of Palearctic Region for Terrestrial Animals, Based on Geographic Distribution of Coleoptera Insects (with a map). Scientific papers of Zoological Institute of Academy of Sciences of USSR, 1935, vol. 2, iss. 2–3, pp. 397–410 (in Russian).
Semenov D. V. Slider Turtle, Trachemys scripta elegans, as Invasion threat (Reptilia; Testudines). Russ. J. of Biological Invasions, 2009, no. 1, pp. 36–44 (in Russian).
Serbinova I. A., Tarkhnishvili D. N. To the Status and Prospects of Conservation of Eastern Spadefoot Toad (Pelobates syriacus) in Georgia. Scientific Studies in Zoo-logical Gardens. Moscow, 2004, iss. 17, pp. 119–126 (in Russian).
Serbinova I. A., Tuniyev B. S. Captivity, Breeding and reintroduction of Northern Banded Newt (Triturus vittatus Jen). Abstracts of First All-Union Meeting on the Issues of the Zooculture. Moscow, 1986, part 2, pp. 147–150 (in Russian).
Serbinova I. A., Shubravyi O. I., Uteshev V. K., Agasyan A. L., Goncharov B. F. Captivity, Breeding and Foundation of a new Natural Population of Eastern Spadefoot Toad (Pelobates syriacus Boettger). Zooculture of Amphibians. Moscow, Publ. House of Academy of Sciences of USSR, 1990, pp. 82–89 (in Russian).
Skorinov D. V., Litvinchuk S. N. Tracing Glacial Refugia of the Smooth Newt (Lissotriton vulgaris) Based on Species Distribution Modelling. Vestnik of Saint Petersburg University, Ser. 3. Biology, 2016, iss. 3, pp. 136–143 (in Russian).
Sobolevssky N. I. A New Form of the Genus Lacerta (Reptilia) from the Crimea. Bull. of Researsh Institute Association at the Physhical-Mathematical Faculty of the First Moscow State University, 1930, vol. 3, no. 2-A, pp. 129–143 (in Russian).
Terentjev P. V., Chernov S. A. A Guid-Book on Reptiles and Amphibians. The 3rd revised edition. Moscow, Soviet Science, 1949. 340 p. (in Russian).
Tertyshnikov M. F. Amphibians of Stavropol Territory. Stavropol’, Stavropol’ State University Press, 1999. 86 p. (in Russian).
Tertyshnikov M. F., Garanin V. I. Anthropogenic Impact on Amphibians and Reptiles and the task of their Protection. Fauna of Stavropol Region. Stavropol’, 1984, iss. 3, pp. 38–48 (in Russian).
Tertyshnikov M. F., Gorovaya V. I. Herpetological Notes. 3. On the reacclimatization of the Rock lizard and the Acclimatization of the Long-legged Wood Frog in the Central Ciscaucasia. Fauna of Stavropol Region. Stavropol’, 1977, iss. 2, pp. 64–69 (in Russian).
Tertyshnikov M. F, Gorovaya V. I. On the Distri-bution and Biology of the Northern Banded newt in the Northern Caucasus. Vestnik Zoologii, 1985, no. 4, pp. 77–79 (in Russian).
Tuniyev B. S. Gerpetofauna gor Al'piiskoi skladchatosti Kavkaza i Srednei Azii: avtoref. dis. … d-ra biol. nauk [Herpetofauna of the Mountains of the Alpine folding of the Caucasus and Central Asia. Thesis of a Dr. Biol. Sciences Diss.]. Saint Petersburg, 1995. 45 p. (in Russian).
Tuniyev B. S., Tuniyev S. B. Herpetofauna of the Sochi National Park. In: Inventory of the Main Taxonomic Groups and Communities, Sozological Studies of the Sochi National Park – the first results of the first National Park in Russia. Moscow, Prestige, 2006, pp. 195–204 (in Russian).
Tuniyev B. S., Tuniyev S. B. European pond turtle (Black Sea population) Emys orbicularis (Linnaeus, 1758). Red Data Book of the Krasnodar Territory (Animals). 2nd edition. Krasnodar, Center for Development of the Krasnodar Territory, 2007, pp. 336–337 (in Russian).
Faizulin A. I., Kukushkin O. V., Ivanov A. Yu., Ermakov O. A. Preliminary Data on the Molecular Genetic Structure of Pelophylax ridibundus (Amphibia: Anura: Ranidae) from the Southern Part of the Crimean Peninsula, Based on Mitochondrial and Nuclear DNA Analysis. Current Studies in Herpetology, 2017, vol. 17, iss. 1–2, pp. 56–65 (in Russian). 
Khlyap L. A., Bobrov V. V., Warshavskiy A. A. Biological Invasions on Russian Territory: Mammals. Russ. J. of Biological Invasions, 2008, vol. 1, no. 2, pp. 78–96 (in Russian).
Khosatzky L. I. About Finding of Tortoises in the European part of the USSR. Priroda, 1948, no. 4, pp. 59–60 (in Russian)
Khonyakina Z. P. Caspian bent-toed Gecko in Dagestan. Priroda, 1965, no. 1, pp. 74 (in Russian).
Tsvelykh A. N. A revision of the Late Pleistocene and Holocene Galliformes faunas of the Crimean Mountains. Zoologicheskii zhurnal, 2016, vol. 95, no. 11, pp. 1354–1361 (in Russian).
Tsemsh I. O. To the systematics and geographical distribution of amphibians and reptiles on Ukraine. Students Scientific Papers of the T. G. Shevchenko Kyiv State Uiversity, 1939, no. 4, pp. 103–117 (in Ukrainian).
Tsurcanu V. F. On the issue of Testudo graeca inhabitation in an interfluve of Dniester and Prut Rivers. Proc. of the First Conference of the Ukrainian Herpetological Society. Kyiv, Zoomuseum NMNH NAS of Ukraine, 2005, pp. 175–179 (in Russian)
Chinchaladze L. M. Caspian bent-toed gecko in Georgia. Priroda, 1956, no. 10, p. 114 (in Russian).
Chkhikvadze V. M. Neogene turtles of the USSR. Tbilisi, Metsniereba, 1989. 102 p. (in Russian).
Chkhikvadze V. M., Bakradze М. А. Whether a chameleon in Western Georgia inhabit?. Reports of the Georgian Academy of Sciences. Tbilisi, 1993, vol. 147, no. 1, pp. 170–173 (in Russian).
Sharygin S. A. Herpetofauna of the Nature Reservation “Cape Martyan”. Scientific Papers of State Nikita Botanical Garden, 1976, vol. 70, pp. 114–120 (in Rus-sian). 
Sharygin S. A. To the study of the rare herpetofaunal species in the Crimea. In: Wildlife Conservation: Abstracts of All-Union Conference of Young Scientists. Moscow, Printing-House of VASKhNIL, 1983, pp. 212–213 (in Russian).
Sharygin S. A. Protection of herpetofauna in the Crimean reserves. In: Theoretical Principles of the Work in Reserves: Abstracts of All-Union Meeting. Moscow, 1985, pp. 304– 307 (in Russian).
Sharygin S. A. Amphibians and Reptiles in the Crimean Reserves. In: Impotrance of Protected Natural Territories in the Biodiversity Consevation: Materials of Conference Dedicated to 75th Anniversary of Kaniv Nature Reserve. Kaniv, Fitosociocentr, 1998, pp. 255–256 (in Russian).
Shelkovnikov A. B. Notes on the amphibians and reptiles of the Aresh county of Elisabethpol province. Re-port on the Caucasian Museum and the Tiflis Public Library for 1908. Tiflis, 1910, pp. 217–242 (in Russian).
Szczerbak N. N. Reptiles as a model for the studying of animals in the process of acclimatization. In: The Problems of Herpetology: Abstracts of Herpetological Conference. Leningrad, Leningrad University Press, 1964, pp. 78–79 (in Russian).
Szczerbak N. N. Zemnovodnye i presmykaiushchiesia Kryma (=Herpetologia Taurica) [Amphibians and Reptiles of the Crimea (=Herpetologia Taurica)]. Kiev, Naukova dumka, 1966 a. 240 p. (in Russian).        
Szczerbak N. N. Amphibians and Reptiles. Simferopol’, Krym, 1966 b. 59 p. (Serie “Nature of the Crimea”) (in Russian). 
Szczerbak N. N. On the Snake Trails. Kiev, Naukova dumka, 1973. 150 p. (in Russian).
Szczerbak N. N. Zemnovodnye i presmykaiushchiesia. Izuchenie fauny i chislennosti nazemnykh pozvonochnykh Karadaga (1981 – 1982) (zakliuchitel'nyi otchet) [Amphibians and Reptiles. The study of Terrestrial Vertebrates Fauna and Abundance at Karadag (1981 – 1982) (final report)]. Letopis' prirody Karadagskogo gosudarstven-nogo zapovednika AN USSR [Annals of Nature of Karadag State Reserve of Academy of Sciences of Ukrainian SSR], 1984, vol. 1, book 1, part 5, pp. 4–32 (in Russian, unpublished).
Szczerbak N. N. Amphibians and Reptiles. In: Fauna of the Karadag State Reserve. Operative-Informational Data. Ed. V. E. Sokolov. Мoscow, VINITI, 1989 а, pp. 33–37 (Serie: Flora and Fauna of the Protected areas of USSR) (in Russian).
Szczerbak N. N. Amphibians and Reptiles. Nature of the Karadag. Kiev, Naukova dumka, 1989 b, pp. 194–197 (in Russian).
Szczerbak N. N., Golubev M. L. Gecko Fauna of the USSR and Adjacent Countries. Kiev, Naukova dumka, 1986. 232 p. (in Russian).
Andreone F., Angelici F. M., Carlino P., Tripepi S., Crottini A. The commom chameleon Chamaeleo chamaeleon in southern Italy: evidence for allochtony of populations in Apulia and Calabria (Reptilia: Squamata: Chamaeleonidae). Italian J. of Zoology, 2016, vol. 83, no. 3, pp. 372–381. 
Arakelyan M. S., Danielyan F. D., Corti C., Sindaco R., Leviton A. E. Herpetofauna of Armenia and Nagorno-Karabakh. Salt Lake City, Society for the Study of Amphibians and Reptiles, 2011. 154 p.
Bauer A. M. How Far North Would the Gecko Move if the Gecko Could Move North?. Herpetological Review, 2000, vol. 31, no. 2, pp. 72–73. 
Bischoff W., Deichsel G. A specimen misidenti-fied as Podarcis muralis (Laurenti, 1768) from Ohio, USA, re-determined as Darevskia valentini (Boettger, 1892) (Reptilia: Lacertidae). Salamandra, 2002, vol. 38, no. 2, pp. 113–117.
Bódis E., Borza P., Potyó I., Puky M., Weiperth A., Guty G. Invasive mollusc, crustacean, fish and reptile species along the Hungarian stretch of the river Danube and some connected waters. Acta Zoologica Academiae Scientarum Hungaricae, 2012, vol. 58, suppl., pp. 29–45.
Böhme W. When does a foreign species deserve a «permit of rezidence»? Non-indigenous spesies (NIS): examples of varying exoticness and varying immigration age, taken from herpetology. Ecology, Ethology and Evolution, 2000, no. 12, pp. 326–328.
Borkin L. J., Litvinchuk S. N., Zuiderwijk A. Bandmolch, Triturus vittatus (Gray, 1835). Handbuch der Reptilien und Amphibien Europas. Bd. 4. Schwan-zlurche II/1. Hrg. K. G. Grossenbacher, B. Thiesmeier. Wiebelsheim, AULA-Verlag, 2003, S. 555–605.
Collins J. T., Gubanyi J. E. History and distribu-tion of the western green lacerta Lacerta bilineata (Reptilia: Squamata: Lacertidae), in Topeka, Kansas. J. of Kansas Herpetology, 2010, vol. 34, pp. 8–9. 
Danielyan F. D., Aslanyan A. On the record of Cas-pian gecko (Tenuidactylus caspius) in Armenia. 12th Ordinary General Meeting of Societas Herpetologica Europaea: Programme and Abstracts. Saint Petersburg, 2003, pp. 53.
Didham R. K., Tylianakis J. M., Hutchison M. A., Ewers R. M., Gemmell N. J. Are invasive species the drivers of ecological change?. Trends in Ecology and Evolution, 2005, vol. 20, no. 9, pp. 470–474.
Dimaki M., Hundsdorfer A. K., Fritz U. Eastern Mediterranean chameleons (Chamaeleo chamaeleon; Ch. africanus) are disticnt. Amphibia–Reptilia, 2008. vol. 29, pp. 535–540.
Dimancea N. Note Upon the Presence of Trachemys scripta elegans (Reptilia) in Oradea City, Romania. Herptologica Romanica, 2013, vol. 7, pp. 41–47.
Domeneghetti D., Marta S., Sbordoni V. A modeling approach for the distribution of two Reptiles species: Mediodactylus kotschyi and Zamenis situla. Atti X Congresso Nazionale Societas Herpetologica Italica. Genova, 2014, pp. 203–205.
Ðorđević S., Anđelković M. Possible reproduction of the Redeared slider, Trachemys scripta elegans (Reptilia: Testudines: Emydidae), in Serbia, under natural conditions. Hyla, 2015, no. 1, pp. 44–49.
Duffus A. L. J., Cunningham A. A. Major disease threats to European amphibians. Herpetological J., 2010, vol. 20, pp. 117–127. 
Erismis U. C., Konuk M., Yoldas T., Agyar P., Yumuk D., Korcan S. E. Survey of Turkey’s endemic amphibians for chytrid fungus Batrachochytridium dendrobatidis. Diseases of Aquatic Organisms, 2014, vol. 111, no. 2, pp. 153–157.
Eversham B. C., Arnold H. R. Introductions and their place in British wildlife. In: Biological Recording of Changes in British Wildlife. Ed. P. T. Harding. London, HMSO, 1992, pp. 44–59.
Ficetola G. F., Scali S. Invasive amphibians and reptiles in Italy. Atti VIII Congresso Nazionale Societas Herpetologica Italica. Pescara, Ianieri Edizioni, 2010, pp. 335–340.
Flärdh B. Herpetofaunan pa Mount Ararat. Snoken, 1983, vol. 13, no. 2, pp. 31–38. 
Fritz U., Ayaz D., Hundsdörfer A. K., Kotenko T., Guicking D., Wink M., Tok C. V., Çiçek K., Buschbom J. Mitochondrial diversity of European pond turtles (Emys orbicularis) in Anatolia and the Ponto-Caspian Region: Multiple old refuges, hotspot of extant diversification and critically endangered endemics. Organisms, Diversity and Evolution, 2009, vol. 9, iss. 2, pp. 100–114.
Fritz U., Guicking D., Lenk P., Joger U., Wink M. When turtle distribution tells European history: mtDNA haplotypes of Emys orbicularis reflect in Germany former division by the Iron Curtain. Biologia, Bratislava, 2004, vol. 59, suppl. 14, pp. 19–25. 
Göçmen B., Veith M., Iğci N., Akman B., Godmann O., Wagner N. No Detection the Amphibian Pathogen Batrachochytridium dendrobatidis in Terrestrial Turkish Salamanders (Lyciasalamandra) Despite its Occurence in Syntopic Frogs (Pelophylax bedriagae). Salamandra, 2013, vol. 49, no. 1, pp. 51–55.
Goldberg S. R., Bursey C. Transport of Helminths to Hawaii via the Brown Anole, Anolis sagrei (Poly-chrotidae). J. of Parasitology, 2000, vol. 86, no. 4, pp. 750–755.
Heym A., Deichsel G., Hochkirch A., Veith M., Schulte U. Do Inrtoduced wall Lizards (Podarcis muralis) cause nishe shifts in a Native Sand lIzard (Lacerta agilis) Population? A case Study from South-Western Germany. Salamandra, 2013, vol. 49, no. 2, pp. 97–104. 
Hoskin C. J. The Invasion and Potential Impact of the Asian House Gecko (Hemidactylus frenatus) in Australia. Austral Ecology, 2011, vol. 36, iss. 3, pp. 240–251.
Kalyabina-Hauf S. A., Deichsel G. Geographic distribution. Lacerta bilineata (Western Green Lizard). Herpetological Review, 2002, vol. 33, no. 3, pp. 225–226.
Krasylenko Yu. A., O. V. Kukushkin O. V. An update of Thin-toed gecko Tenuidactylus bogdanovi (Reptilia: Gekkonidae) population status in Odessa City, Ukraine. Proc. of the Zoological Museum, Kyiv, 2017, no. 48, pp. 3 – 12. 
Krysko K. L., Burgess J. P., Rochford M. R., Gilette C. R., Cueva D., Enge K. M., Somma L. A., Stabile J. L., Smith D. C., Wasilewski J. A., Kiechhefer G. N., Granatovsky M. C., Nielsen S. V. Verified non-indigenous amphibians and reptiles in Florida from 1863 through 2010: Outlining the invasion process and identifying invasion pathways and stages. Zootaxa, 2011, vol. 3028, pp. 1–64. 
Kukushkin O., Jablonski D. A record of the Balkan Stripe-necked terrapin, Mauremys rivulata (Testudines: Geoemydidae) from the Azov Sea Coast in the Crimea. Amphibian and Reptile Conservation, 2016, vol. 10, no. 2, pp. 27–29.
Kyriazi P., Kornilios P., Nagy Z. T., Poulakakis N., Kumlutaş Y., Ilgaz Ç., Avci A., Göçmen B., Lymberakis P. Comparative phylogeography reveals distinct colonization patterns of Cretan snakes. J. of Biogeography, 2013, vol. 40, iss. 6, pp. 1143–1155.
Langton T. E. S., Atkins W., Herbert C. On the distribution, ecology and management of nonnative rep-tiles and amphibians in the London Area. Part 1. Distri-bution and predator / prey impacts. London Naturalist, 2011, no. 90, pp. 83–156.
Lever C. Naturalized Reptiles and Amphibians of the World. Oxford, New York, Oxford University Press, 2003. 318 p.
Lymberakis P., Poulakakis N. Three Continents Claiming an Archipelago: The Evolution of Aegean’s Herpetofaunal Diversity. Diversity, 2010, vol. 2, iss. 2, pp. 233–255. 
Majláthova V., Majláth I., Hromada M., Tryjanowski P., Bona M., Antczak M., Víchová B., Dzimko Š., Michalca A., Pet’ko B. The role of the sand lizard (Lacerta agilis) in the transmission cycle of Borrelia burgdorfei sensu lato. Intern. J. of Medical Microbiology, 2008, vol. 298, suppl. 1, pp. 161–167. 
Marzahn E., Mayer W., Joger U., Ilgaz Ç., Jablon-ski D., Kindler C., Kumlutaş U., Nistri A., Schneeweiss N., Vamberger M., Žagar A., Fritz U. Phylogeography of the Lacerta viridis complex: mitochondrial and nuclear markers provide taxonomic insights. J. of Zoological Sys-tematics and Evolutionary Research, 2016, vol. 54, iss. 2, pp. 85–105. 
Mateo J. A., Ayres C., López-Jurado L. F. Los Anfibios y Reptiles Naturalizados en España: Historia y Evolución de una Problemática Crecienta. Boletín de la Asociación Herpetológica Española, 2011, vol. 22, pp. 2–42.
Merzlikin I. R. Finds of the Red-eared slider Trachemys scripta elegans (Reptilia, Testudines) in the Natural Biotopes of Ukraine. The IV Intern. Symp. Invasion of alien species in Holarctic. Yaroslavl’, Filigran’, 2013, p. 118. 
Mulder J. Herpetological Observation in Turkey (1987 – 1995). Deinsea (Annual of the Natural History Museum in Rotterdam), 1995, no. 2, pp. 51–66.
National Invasive Species Council (NISC). Invasive Species Definition Clarification and Guidance Wait Paper. Submitted by the Definitions Subcommittee of the Invasive Species Advisory Committee (ISAC). Meet of In-vasive Species Challenge. Washington, 2006. 11 p.
Nekrasova O. D., Kostiushyn V. A. Current Distribution of the Introduced Rock Lizards of the Darevskia (saxicola) complex (Sauria, Lacertidae, Darevskia) in Zhytomyr Region (Ukraine). Vestnik Zoologii, 2016, vol. 50, no. 3, pp. 225–230. 
Pallas P. S. Zoographia rosso-asiatica, sistens omnium animalium in extenso imperio Rossico et adjacentibus maribus observatorum recensionem, domicilia, mores et descriptiones anatomen atque icones plurimorum. Petropoli, Academiae Scientarum Impress, 1831, vol. 3. Animalia monocardia seu frigidi sanguinis. 549 p. 
Pérez-Santigosa N., Florencio M., Hidalgo-Vila J., Díaz-Paniagua C. Does the Exotic Turtle, Trachemys scripta elegans, Compete for Food With Coexisting Native Turtles?. Amphibia – Reptilia, 2011, vol. 32, iss. 2, pp. 167–175.
Pinya S., Carretero M. The Balearic herpetofauna: а species update and a review on the evidence. Acta Herpetologica, 2011, vol. 6, no. 1, pp. 59–80.
Poulakakis N., Kapli P., Kardamaki A., Skour-tanioti E., Gösmen B., Ilgaz Ç., Kumlutaş Y., Avci A., Lymberakis P. Comparative Phylogeography of Six Herpetofauna Species in Cyprus : Late Miocene tо Pleistocene Colonization Routes. Biological J. of the Linnean Society, 2013, vol. 108, iss. 3, pp. 619–635.
Scalera R. Trachemys scripta. Datasheet DAISIE (Delivering Alien Invasive Species Inventories for Europe). 2006. Available at: Trachemys_scripta.pdf (accessed 30 May 2017). 
Schulte U., Veith M., Hochkirch A. Rapid Genetic Assimilation on Native Wall Lizard Population (Podarcis muralis) Through Extensive Hybridization With Intro-duced Lineages. Molecular Ecology, 2012, vol. 21, iss. 17, pp. 4313–4326. 
Semyenova S. K., Korsunenko A. V., Vasilyev V. A., Pereschkolnik S. L., Mazanaeva V. A., Bannikova A. A., Ryskov A. P. RAPD Variation in Mediterranean Turtle Testudo graeca (L.) (Testudinidae). Russ. J. of Genetics, 2004, vol. 40, no. 12, pp. 1348–1355. 
Silva-Rocha I., Salvi D., Carretero M. A. Genetic data reveal a multiple origin for the populations of the Italian wall lizard Podarcis sicula (Squamata: Lacertidae) introduced in the Iberian Peninsula and Balearic islands. Italian J. of Zoology, 2012, vol. 79, no. 4, pp. 502–510. 
Široký P., Bĕlohlávek T., Papoušek I., Jandzik D., Miculíček P., Kubelová M., Zdražilová-Dubská L. Hidden Threat of Tortoise Ticks: High Prevalence of Crimean-Congo Haemorrhagic Fever Virus in the Ticks Hyalomma aegyptium in the Middle East. Parasites and Vectors, 2014, vol. 7, pp. e101. DOI: 10.1186/1756-3305-7-101
Speth J. D., Tchernov E. Middle Paleolithic tortoise use at Cebara cave (Israel). J. of Archaeological Science, 2002, vol. 29, no. 5, pp. 471–483.
Stöck M., Grifoni G., Armor N., Scheidt U., Sicilia A., Novarini N. On the Origin of the Recent HerpetoFauna of Sicily: Comparative Phylogeography Using Homologous Mitochondrial and Nuclear Genes. Zoologischer Anzeiger, 2016, vol. 261, pp. 70–86.
Tarkhnishvili D., Gabelaia M., Kandauriv A., Bukhnikashvili A., Iankoshvili G. Isolated Population of the Middle Eastern Phoenicolacerta laevis from the Georgian Black Sea Coast, and its Genetic Closeness to Populations from Southern Turkey. Zoology in the Middle East, 2017, vol. 63, iss. 4, pp. 311–315. 
Urošević A., Tomović L., Ajtić R., Simović A., Džukić G. Alterations in the reptilian fauna of Serbia : In-troduction of exotic and anthropogenic range expansion of native species. Herpetozoa, 2016, vol. 28, no. 3–4, pp. 115–132. 
Vamberger M., Lipovšek G. First reproduction record of Trachemys scripta (Schoepff, 1792), in Slovenia. Hepetozoa, 2012, vol. 25, no. 1–2, pp. 76–79.
Wirga M., Majtyka T. Do Climate Requirements Explain the Northern Range of European Reptiles? Common wall lizard Podarcis muralis (Laur.) (Squamata, Lacertidae) as an example. North-Western J. of Zoology, 2015, vol. 11, no. 2, pp. 296–303.
Witmer G. W., Fuller P. L. Vertebrate Species Introductions in the United States and its Territories. Current Zoology, 2011, vol. 57, no. 5, pp. 559 – 567. 
Zenni R. D., Nuñez M. A. The Elephant in the Room: the Role of Failed Invasions in Understanding Invasion Biology. Oikos, 2013, vol. 122, no. 6, pp. 801–815.

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